Review of the gobionids of Iran (Family Gobionidae)

Brian W. COAD


The systematics, morphology, distribution, biology and economic importance of the gobionids of Iran are described, the species are illustrated, and a bibliography on these fishes in Iran is provided. There are three native species in the genera Gobio and Romanogobio found in northeastern and northwestern Iran respectively and a widely introduced exotic species Pseudorasbora parva.


Biology, Morphology, Exotic, Abbottina, Gobio, Romanogobio, Pseudorasbora.

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Aazami, J.; Esmaili-Sari, A.; Abdoli, A.; Sohrabi, H. & Van den Brink, P.J. 2015. Length-weight relationship of 14 fish species from Tajan River, southern Caspian Sea basin, Iran. Iranian Journal of Ichthyology 2(4): 299-301.

Abbasi, K. 2009. Studying native fishes in Hamadan province (Iran). Agricultural Research and Education Organization, Iranian Fisheries Research Institute, Tehran. MS Report. 165 p. (In Farsi)

Abbasi, K.; Khodaparast, H.; Moradi, M. & Sarpanah, A.N. 2016. Identification and abundance of fishes in Kardeh Reservoir (Mashad, Iran). The Fourth Iranian Conference of Ichthyology, Ferdowsi University of Mashhad, 20-21 July 2016 (abstract).

Abbasi, K.; Moradi, M.; Nikpoor, M.; Zahmatkesh, Y.; Mirzajani, A.; Mousavi-Sabet, H.; Abdoli, A.; Sayadrahim, M.; Mohammadidost, R.; Hoseinjani, A.; Zolfinezhad, K. & Ramzani, M. 2017. Study on distribution of fish species in Anzali Wetland and its rivers. The Fifth Iranian Conference of Ichthyology, Islamic Azad University of Babol, 13-14 December 2017 (abstract and p. 642-648. In Farsi).

Abbasi, K.; Moradkhah, S. & Sarpanah, A.N. 2007. Identification and distribution of fish fauna in Siahdarvishan River (Anzali Wetland basin). Pajouhesh va Sazandegi 19(1)(74): 27-39. (In Farsi)

Abbasi, K.; Nikseresht, K. & Noroozi, H. 2009. Identification of fish population in Agh-Gol, Pir-Salman, Gamasiab and Haram-abad wetlands of the Hamadan Province. Journal of Wetland Ecobiology 1(1): 71-90. (In Farsi)

Abbasi, K.; Salavatian, S.M. & Abdollapoor, H. 2005. Investigating fish diversity and distribution in the Mahabad-Chai River of the Lake Urmia basin, north-western Iran. Iranian Journal of Fisheries Sciences 13(4): 75-94. (In Farsi)

Abbasi, K.; Valipour, A.; Talebi Haghighi, D.; Sarpanah, A. & Nezami, Sh. 1999. Atlas of Iranian Fishes. Gilan Inland Waters. Gilan Fisheries Research Centre, Rasht. 113 p. (In Farsi)

Abdoli, A. 1992. A new fish species in Mazandaran ab bandans. Abzeeyan, Tehran 2(8): 54-55. (In Farsi)

Abdoli, A. 2000. The Inland Water Fishes of Iran. Iranian Museum of Nature and Wildlife, Tehran. 378 p. (In Farsi)

Abdoli, A. & Naderi, M. 2009. Biodiversity of Fishes of the Southern Basin of the Caspian Sea. Abzian Scientific Publications, Tehran. 243 p. (In Farsi)

Abdoli, A.; Naderi, M.; Foroughifrad, H. & Kiabi, B.H. 2014. Fish diversity and distribution in two protected rivers, Sardabrud and Chalus, southern Caspian Sea basin, Iran. Iranian Journal of Ichthyology 1(2): 91-95.

Abdurakhmanov, Yu.A. 1962. Ryby Presnykh vod Azerbaidzhana [Freshwater Fishes of Azerbaidzhan]. Akademii Nauk Azerbaidzhanskoi SSR, Institut Zoologii, Baku. 407 p. Aliev, D. S., Sukhanova, A. I. & Shakirova, F. M. 1988. Ryby vnutrennikh vodoemov Turkmenistana [Fishes of the inland waters of Turkmenistan]. Ylym, Ashkhabad. 142 p.

Aliev, D.S.; Sukhanova, A.I. & Shakirova, F.M. 1988. Ryby vnutrennikh vodoemov Turkmenistana [Fishes of the inland waters of Turkmenistan]. Ylym, Ashkhabad. 142 p.

Alizadeh Marzenaki, A.; Shojaei Kavan, L.; Taghiyan, H. & Shahriari, R. 2016. A survey of biodiversity and distribution of fish in Gavehrud of Kermanshah Province. Breeding and Aquaculture Sciences Quarterly 3(8): 69-84. (In Farsi)

Amanov, A.A. 1972. Morphology and mode of life of the Turkestan gudgeon [Gobio gobio lepidolaemus (Kessler)] from the Surkhandar'ya Basin. Journal of Ichthyology 12(4): 601-608.

Anonymous. 1994. Report of a new cyprinid fish in Gilan rivers. Scientific Newsletter of the Iranian Fisheries Research and Training Organization p. 12-13. (In Farsi)

Arai, R. 2011. Fish Karyotypes. A Check List. Springer, Tokyo. 340 p.

Asadi, H.; Sattari, M.; Motalebi, Y.; Zamani-Faradonbeh, M. & Gheytasi, A. 2017. Length-weight relationship and condition factor of seven fish species from Shahrbijar River, southern Caspian Sea basin, Iran. Iranian Journal of Fisheries Sciences 16(2): 733-741.

Asgardun, S. & Nowferesti, H. 2014. Length-weight relationship of Pseudoras (sic) parva from the Mashhad Chalidare Dam, Iran. The Second Iranian Conference of Ichthyology, Faculty of Natural Resources, University of Tehran, Karaj, 7-8 May 2014 (abstract).

Banarescu, P. 1961. Weitere systematische Studien über die Gattung Gobio (Pisces, Cyprinidae) insbesondere im Danaubecken. Vĕstník československé Společnosti zoologické 25: 318-416.

Bănărescu, P.M. 1992. A critical updated checklist of Gobioninae (Pisces, Cyprinidae). Travaux du Muséum d'Histoire naturelle "Grigore Antipa", Bucarest 32: 303-330.

Bănărescu, P.M. 1999. The Freshwater Fishes of Europe. Volume 5. Cyprinidae 2. Part I: Rhodeus to Capoeta. AULA-Verlag, Wiebelsheim. 427 p.

Banarescu, P. & Nalbant, T. 1973. Pisces, Teleostei, Cyprinidae (Gobioninae). Das Tierreich, Berlin, 93. 304 p.

Bănărescu, P.M. & Paepke, H-J. 2002. The Freshwater Fishes of Europe. Volume 5/III. Cyprinidae 2. Part III: Carassius to Cyprinus. Gasterosteidae. AULA-Verlag, Wiebelsheim. 305 p.

Barzegar, M. & Jalali, B. 2002. Parasites of Kaftar Lake fishes, their geographical distribution and economical importance. Science Journal of the School of Veterinary Medicine, Shahid Chamran University, Ahvaz 3(5): 52-64. (In Farsi)

Barzegar, M. & Jalali, B. 2009. Crustacean parasites of fresh and brackish (Caspian Sea) water fishes of Iran. Journal of Agricultural Science and Technology 11: 161-171.

Barzegar, M. & Jalali Jafari, B. 2006. Helminthes, Acanthocephala and Crustacean parasites of fishes in Vahdat Reservoir. Iranian Journal of Veterinary Science 3: 229-234. (In Farsi)

Berg, L.S. 1932. Ryby presnykh vod SSSR i sopredel’nykh stran [Freshwater fishes of the USSR and adjacent countries]. 3rd Edition, Part 1. Vsesoyuznyy Institut Ozernogo i Rechnogo Rybnogo Khozyaistva, Leningrad. 543 p.

Berg, L.S. 1948-1949. Freshwater fishes of the USSR and adjacent countries. Israel Program for Scientific Translations, Jerusalem (1962-1965). 3 volumes.

Biukani, S.; Safarpour Amlashi, A. & Falahatkar, B. 2013. Fish fauna of Gamasiab River in Kermanshah Province. Journal of Fisheries Science and Technology 2(1): 1-12. (In Farsi)

Boltachev, A.R.; Danilyuk, O.N.; Pakhorukov, N.P. & Bondarev, V.A. 2006. Distribution and certain features of the morphology and biology of the stone moroko Pseudorasbora parva (Cypriniformes, Cyprinidae) in the waters of Crimea. Journal of Ichthyology 46(1): 58-63.

Bozorgnia, A.; Youssefi, M.R.; Barzegar, M.; Hosseinifard, S.M. & Ebrahimpour, S. 2012. Biodiversity of parasites of fishes in Gheshlagh (Vahdat) Reservoir, Kurdistan Province, Iran. World Journal of Fish and Marine Sciences 4(3): 249-253.

Coad, B.W. 1996. Exotic fish species in the Tigris-Euphrates basin. Zoology in the Middle East 13: 71-83.

Coad, B.W. & Abdoli, A. 1993. Exotic fish species in the fresh waters of Iran. Zoology in the Middle East 9: 65-80.

Courtenay, W.R. & Stauffer, J.R. 1984. Distribution, Biology, and Management of Exotic Fishes. Johns Hopkins University Press, Baltimore. 430 p.

Eagderi, S. & Moradi, M. 2017. Range extension of the lake goby Rhinogobius similis Gill, 1859 (Teleost (sic): Gobiidae) to Urmia Lake basin in northwestern Iran. Biharean Biologist 11(2): 123-125.

Eagderi, S. & Nasri, M. 2012. A first record of the Bittrling (sic) Rhodeus amarus (Bloch, 1782) Cypriniformes, Cyprinidae) in the Iranian part of Tigris-Euphrates Basin. International Research Journal of Applied and Basic Sciences 3(3): 639-641.

Eschmeyer, W.N. 1990. Catalog of the Genera of Recent Fishes. California Academy of Sciences, San Francisco. 697 p.

Eschmeyer, W.N.; Ferraris, C.J.; Hoang, M.D. & Long, D.J. 1996. A Catalog of the Species of Fishes. Preliminary version (Sept. 1996).

Esmaeili, H.R.; Babai, S.; Gholamifard, A.; Pazira, A.; Gholamhosseini, A. & Coad, B.W. 2015. Fishes of the Persis region of Iran: an updated checklist and ichthyogeography. Iranian Journal of Ichthyology 2(3): 201-223.

Esmaeili, H.R. & Ebrahimi, M. 2006. Length-weight relationships of some freshwater fishes of Iran. Journal of Applied Ichthyology 22: 328-329.

Esmaeili, H.R.; Gholamifard, A. & Gholamhosseini, A. 2010. Ichthyofauna of Iranian part of Tigris River basin. 16th National and 4th International Conference of Biology, Ferdowsi University of Mashhad, Mashhad, Iran, 14-16 September 2010 (abstract).

Esmaeili, H.R.; Malekzehi, H.; Pazira, A. & Freyhof, J. 2013a. First record of the Kalabans, Bangana dero (Hamilton, 1822), from Iran (Teleostei: Cyprinidae). Zoology in the Middle East 59(1): 89-91.

Esmaeili, H.R.; Malekzehi, H.; Pazira, A. & Freyhof, J. 2013b. First record of the Kalabans, (Cyprinidae: Bangana dero), from Iran. The First Iranian Conference of Ichthyology, Isfahan University of Technology, 15-16 May 2013, p. 8 (abstract).

Esmaeili, H.R.; Sayyadzadeh, G.; Eagderi, S. & Abbasi, K. (2018). Checklist of freshwater fishes of Iran. FishTaxa 3(3): 1-95.

Esmaeili, H.R., Teimori, A. & Abdoli, A. 2015. Fish Diversity of Fars Province. MS, 257 p. (In Farsi)

Fricke, R., Bilecenoglu, M. & Sari, H. M. 2007. Annotated checklist of fish and lamprey species (Gnathostomata and Petromyzontomorphi) of Turkey, including a Red List of threatened and declining species. Stuttgarter Beiträge zur Naturkunde, Serie A (Biologie) 706: 169 p.

Ghasemi, H.; Jouladeh Roudbar, A.; Eagderi, S.; Abbasi, K., Vatandoust, S. & Esmaeili, H.R. 2015. Ichthyofauna of Urmia basin: Taxonomic diversity, distribution and conservation. Iranian Journal of Ichthyology 2(3): 177-193.

Gholizadeh, M.; Khosozadeh, F. & Peyvandi, N. 2014. Study on faunistic of fish fauna in Kelarud River (Babol), (Mazandaran province). Journal of Applied Ichthyological Research 2(1): 1-10. (In Farsi).

Gozlan, R.E.; Andreou, D.; Asaeda, T.; Beyer, K.; Bouhadad, R.; Burnard, D.; Caiola, N.; Cakic, P.; Djikanovic, V.; Esmaeili, H.R.; Falka, I.; Golicher, D.; Harka, A.; Jeney, G.; Kováč, V., Musil, J.; Nocita, A.; Povz, M.; Poulet, N., Virbickas, T.; Wolter, C.; Tarkan, A.S.; Tricario, E.; Trichkova, T.; Verreycken, H.; Witkowski, A.; Zhang, C.; Zweimueller, I. & Britton, J.R. 2010. Pan-continental invasion of Pseudorasbora parva: towards a better understanding of freshwater fish invasions. Fish and Fisheries 11(4): 315-340.

Günther, R.T. 1899. Contributions to the natural history of Lake Urmi, N.W. Persia, and its neighbourhood. Journal of Zoology of the Linnaean Society 27: 345-453, pl. 21-30.

Hardouin, E.A.; Andreou, D.; Zhao, Y.; Chevret, P.; Fletcher, D.H., Britton, J.R. & Gozlan, R. E. 2018. Reconciling the biogeography of an invader through recent and historic genetic patterns: the case of topmouth gudgeon Pseudorasbora parva. Biological Invasions 20(8): 2157-2171.

Hasankhani, M.; Keivany, Y.; Daliri, M.; Pouladi, M. & Soofiani, N. M. 2014. Length-weight and length-length relationships of four species (Barbus lacerta Heckel, 1843), Oxynoemacheilus angorae (Steindachner, 1897), Squalius lepidus (Heckel, 1843) and Pseudorasbora parva (Temminck & Schlegel, 1846) from the Sirwan River (western Iran). Journal of Applied Ichthyology 30(1): 206-207.

Hosseini, S.A.H.; Mirvaghefi, A.R.; Zare, P. & Nasiri, E. 2011. Investigation of infection by plerocercoid stage of Ligula intestinalis in amorcheh gastric “Pseudorasbora parva”, in Zabol Chahnimeh reservoirs. Journal of Fisheries 5(2)(18): 141-148.

Hosseini Fard, S.M.; Talebi, S.A. & Alaedini Shourmasti, A. 2017. Investigation of the amount of parasitic fauna of waterlogged fish in Langour in Babol. The Fifth Iranian Conference of Ichthyology, Islamic Azad University of Babol, 13-14 December 2017, p. 817-821. (In Farsi)

IUCN. 2015. The IUCN Red List of Threatened Species. Version 2015.2. International Union for the Conservation of Nature, Gland, Switzerland (

Jackson, M.C. & Britton, J. R. 2013. Variation in the trophic overlap of invasive Pseudorasbora parva and sympatric cyprinid fishes. Ecology of Freshwater Fish 22(4): 654-657.

Jolodar, M.N. & Abdoli, A. 2004. Fish Species Atlas of South Caspian Sea Basin (Iranian Waters). Iranian Fisheries Research Organization, Teheran.110 p. (In Farsi and English)

Jouladeh Roudbar, A.; Eagderi, S. & Esmaeili, H.R. 2015. Fishes of the Dasht-e Kavir basin of Iran: an updated checklist. International Journal of Aquatic Biology 3(4): 263-273.

Kamensky, S.N. 1899-1901. Karpovye (Cyprinidae) Kavkaza i Zakavkaz'ya [Die Cypriniden der Kaukasusländer und ihrer angrenzenden Meere]. Tiflis. 157 p., 192 p. (In Russian and German)

Karimpour, M. 1998. The ichthyofauna of Anzali Lagoon. Iranian Journal of Fisheries Sciences 7(2): 83-94, 9. (In Farsi)

Khaval, A.; Abbasi, K. & Valipour, A. 2010. Biological role of northern pike, Esox lucius (Linnaeus 1758) in control of aquatic pests in carps culture ponds. Iranian Scientific Fisheries Journal 19(2): 39-50.

Khoshnood, Z. 2014. Identification and study of fish species in Karkheh River (Iran). Transylvanian Review of Systematical and Ecological Research 16(2): 97-106.

Khosravi Katuli, K.; Eagderi, S. & Nowferesti, H. 2013. Phenotypic plasticity of body shape in three populations of false rasbora (Pseudorasbora parva) using geometric morphometric method. Journal of Utilization and Cultivation of Aquatics 2(4): 61-71. In Farsi.

Kiabi, B.H.; Abdoli, A. & Naderi, M. 1999. Status of the fish fauna in the South Caspian Basin of Iran. Zoology in the Middle East 18: 57-65.

Klinkhardt, M.; Tesche, M. & Greven, H. 1995. Database of Fish Chromosomes. Westarp Wissenschaften, Magdeburg. 237 p.

Kottelat, M. & Freyhof, J. 2007. Handbook of European Freshwater Fishes. Kottelat, Cornol, Switzerland and Freyhof, Berlin, Germany. 646 p.

Makeyeva, A.P. & Mokhamed, M.I.Z. 1982. The reproduction and development of Pseudorasbora parva in Soviet Central Asia. Journal of Ichthyology 22(1): 69-82.

Malek, M. & Mobedi, I. 2001. Occurrence of Clinostomum Complanatum (sic) (Rudolphi, 1819) (Digenea: Clinostomatidae) in Capoeta capoeta gracilis (Osteichthys (sic): Cyprinidae) from Shiroud River, Iran. Iranian Journal of Public Health 30(3-4): 95-98.

Malekzehi, M.H.; Esmaeili, H.R.; Zareian, H.; Farahani, Z. & Pazira, A. 2014. Incidence of Lernaea (Crustacea: Copepoda) parasitic in the Maskhkid River basin, southeast of Iran. International Journal of Aquatic Biology 2(1): 9-13.

Mendel, J.; Lusk, S.; Vasil’eva, E.D.; Vasil’ev, V.P.; Lusková, V.; Guler Ekmekci, F.; Erk’akan, F.; Ruchin, A.; Koščo, J.; Vetešník, L.; Halačka, K.; Šanda, R.; Pashkov, A.N. & Reshetnikov, S.I. 2008. Molecular phylogeny of the genus Gobio Cuvier, 1816 (Teleostei: Cyprinidae) and its contribution to taxonomy. Molecular Phylogenetics and Evolution 47(3): 1061-1075.

Moradi, M. & Eagderi, S. 2014. First record of the lake goby Rhinogobius similis Gill, 1859 from the Urmia Lake basin, Iran. Biyoloji Bilimleri Araştırma Dergisi 7(2): 52-55.

Mousavi-Sabet, H.; Ganjbakhsh, B.; Geiger, M.F. & Freyhof, G. 2016. Redescription of Gobio nigrescens from the Hari River drainage (Teleostei: Cyprinidae). Zootaxa 4114(1): 71-80.

Movchan, Yu.V. & Kozlov, V.I. 1978. Morphological characters and certain ecological characteristics of the Chinese chebachok Pseudorasbora parva in Ukrainian reservoirs. Hydrobiological Journal 14(5): 36-41.

Naseka, A.M. 1996. Comparative study on the vertebral column in the Gobioninae (Cyprinidae, Pisces) with special reference to its systematics. Publicaciones Especiales Instituto Español de Oceanografía 21: 149-167.

Naseka, A.M. & Freyhof, J. 2004. Romanogobio parvus, a new gudgeon from River Kuban, southern Russia (Cyprinidae, Gobioninae). Ichthyological Exploration of Freshwaters 15(1): 17-23.

Patimar R. 2008. Fish species diversity in the lakes of Alma-Gol, Adji-Gol, and Ala-Gol, Golestan Province, northern Iran. Journal of Ichthyology 48(10): 911-917.

Patimar, R. & Baensaf, S. 2012. Morphology, growth and reproduction of the non-indigenous topmouth gudgeon Pseudorasbora parva (Temminck et Schlegel, 1846) in the wetland of Alma-Gol, northern Iran. Russian Journal of Biological Invasions 3(1): 71-75.

Patimar, R.; Igdrei, S.; Hamid-Sales, B. & Ba-Ensaf, S. 2002a. Two alien species in the International Wetlands, Alma-Gol, Adji-Gol and Ala-Gol, of Golestan Province, Iran. World Congress on Aquatic Protected Areas, Cairns, 14-17 August 2002 (abstract).

Patimar, R.; Igdrei, S.; Hamid-Sales, B. & Ba-Ensaf, S. 2002b. Two alien species in the International Wetlands, Alma-Gol, Adji-Gol and Ala-Gol, of Golestan Province, Iran (a new record). Australian Society for Fishery Biology Annual Conference, 2002 (abstract).

Pipoyan, S.Kh. 1996. Pseudorasbora parva (Cypriniformes, Cyprinidae) in water bodies of the Ararat Plain (Armenia). Journal of Ichthyology 36(7): 536-538.

Pipoyan, S.Kh. 1998. Gobio gobio (L.) (Cyprinidae), a new species of fauna in Armenia. Journal of Ichthyology 38(6): 441-446.

Radkhah, A. & Eagderi, S. 2015. Length-weight and length-length relationships and condition factor of six cyprinid species from Zarrineh River (Urmia Lake basin, Iran). Iranian Journal of Ichthyology 2(1): 61-64.

Rahimi, S. & Tabiee, O. 2013. Analysis of fish faunas in Kaftar Wetlands: A case study of Fars Province (Iran). International Journal of Agriculture and Crop Sciences 6(6): 338-341.

Ramin, M.; Bagheri, S.; Moradi, M.; Abbasi, K.; Mirzajani, A. & Doustdar, M. 2016. Identification, distribution, abundance and species composition of fish in the Chitgar Lake (Tehran Province of Iran). Iranian Journal of Fisheries Sciences 15(1): 585-589.

Ramin, M. & Doustdar, M. 2017. Introducing and determination abundance of invasive and alien fishes of Chitgar Lake. The Fifth Iranian Conference of Ichthyology, Islamic Azad University of Babol 13-14 December 2017, p. 722-727. (In Farsi)

Reshetnikov, Yu.S.; Bogutskaya, N.G.; Vasil'eva, E.D.; Dorofeeva, E.A.; Naseka, A.M.; Popova, O.A.; Savvaitova, K.A.; Sideleva, V.G. & Sokolov, L.I. 1997. An annotated check-list of the freshwater fishes of Russia. Journal of Ichthyology 37(9): 687-736.

Reshetnikov, Yu.S. & Shakirova, F.M. 1993. Zoogeograficheskii analiz ikhtiofauny srednei Asii po spiskam presnovodnykh ryb [Zoogeographical analysis of the ichthyofauna of Central Asia by lists of freshwater fish]. Voprosy Ikhtiologii 33(1): 37-45.

Rezaei Tavabe, K. & Azarnivand, H. 2013. Biodiversity in qanats (the case study of Kerman County, Iran). Desert 18: 99-104.

Robins, C.R.; Bailey, R.M.; Bond, C.E.; Brooker, J.R.; Lachner, E.A.; Lea, R.N. & Scott, W.B. 1991. World Fishes Important to North Americans Exclusive of Species from the Continental Waters of the United States and Canada. American Fisheries Society Special Publication 21. 243 p.

Salavatian, S.M.; Abbasi, K. & Abedini, A. 2016. Identification, abundance and length, weight and age composition of fishes in Arasbaran Reservoir. The Fourth Iranian Conference of Ichthyology, Ferdowsi University of Mashhad, 20-21 July 2016 (abstract).

Sal'nikov, V.B. 1995. Possible changes in the composition of the ichthyofauna after completion of the Karakum Canal in Turkmenistan. Journal of Ichthyology 35(7): 108-121.

Sal'nikov, V.B. 1998. Anthropogenic migration of fish in Turkmenistan. Journal of Ichthyology 38(8): 591-602.

Shakirova, F.M. & Sukhanova, A. I. 1994. Iktiofauna Turkmenistana (sostav i rasprostranenie) [Ichthyofauna of Turkmenistan (composition and distribution)]. Izvestiya Akademii Nauk Turkmenistana, Seriya Biologicheskikh Nauk 3(1993): 35-45.

Stout, C.C.; Tan, M.; Lemmon, A.R.; Lemmon, E.M. & Armbruster, J.W. 2016. Resolving Cypriniformes relationships using an anchored enrichment approach. BMC Evolutionary Biology 16: 244.

Svetovidova, A.A. 1978. [List of holotypes, syntypes and paratypes kept in the Department of Ichthyology of the Moscow State University Museum]. Sbornik Trudov Zoologicheskogo Muzeya MGU (Archives of the Zoological Museum of Moscow State University) 16: 256-263.

Taghiyan, H.; Sahapouri, M.; Mohammadi, V. & Alizadeh, A. 2016. A survey on biodiversity, abundance and distriubution of Dinorab River fish. Breeding and Aquaculture Sciences Quarterly 3(7): 29-42. (In Farsi)

Tan, M. & Armbruster, J.W. 2018. Phylogenetic classification of extant genera of fishes of the order Cypriniformes (Teleostei: Ostariophysi). Zootaxa 4476(1): 6-39.

Tang, K.L.; Agnew, M.K.; Chen, W-J.; Hirt, M.V.; Raley, M.E.; Sado, T.; Schneide, L.M.; Yang, L.; Bart, H.L.; He, S.; Liu, H.; Miya, M.; Saitoh, K.; Simons, A.M.; Wood, R.M. & Mayden, R.L. 2011. Phylogeny of the gudgeons (Teleostei: Cyprinidae: Gobioninae). Molecular Phylogenetics and Evolution 61(1): 103-124.

Teimori, A.; Esmaeili, H.R. & Gholamhosseini, A. 2010. The ichthyofauna of Kor and Helleh River basins in southwest of Iran with reference to taxonomic and zoogeographic features of native fishes. Iranian Journal of Animal Biosystematics 6(1): 1-8.

Wildekamp, R.H.; Van Neer, W.; Küçük, F. & Ünlüsayin, M. 1997. First record of the eastern Asiatic gobionid fish Pseudorasbora parva from the Asiatic part of Turkey. Journal of Fish Biology 51(4): 858-861.

Záhorská, E.; Kováč, V.; Falka, I.; Beyer, K.; Katina, S.; Copp, G.H. & Gozlan, R.E. 2009. Morphological variability of the Asiatic cyprinid, topmouth gudgeon Pseudorasbora parva, in its introduced European range. Journal of Fish Biology 74(1): 167-185.

Zamani Faradonbe, M.; Eagderi, S. & Poorbagher, H. 2017. Niche overlap in fish assemblages inferred from canonical correspondence analysis: A case study with the Totkabon River, north of Iran. Ege Journal of Fisheries and Aquatic Sciences 34(2): 151-156.

Zhao, J.; Xu, D.; Zhao, K.; Diogo, R.; Yang, J. & Peng, Z. 2016. The origin and divergence of Gobionine fishes (Teleostei: Cyprinidae) based on complete mitochondrial genome sequences. Journal of Applied Ichthyology 32(1): 32-39.


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